SDH provides a way for plans to convert sorbitol into fructose without using ATP. According to the reports of Bianco et al. and Li and Li, the increase of SDH activity was a key factor for catabolism of sorbitol in response to Nutlin-3 drought stress. The sdh-mutants of Arabidopsis with inhibition of SDH activity exhibited less dry weight and root length compared to wild-type under supply of exogenous sorbitol condition. Therefore, it can be deduced that increased SDH activity induced by exogenous Spm in both white clover cultivars may play a part in converting sorbitol into fructose to keep balance for metabolism of sorbitol. Under abiotic stress, exogenous ABA-, cytokinin- or proline-induced increase in the expression of dehydrins proves that various phytohormones or physiological activators are associated with regulation of dehydrins in plants. Previous studies have indicated that Spm and dehydrins had the same functions of scavenging reactive oxygen species and maintaining the structure of membrane. Although it was confirmed that Spm and dehydrins both could enhance the drought tolerance of plants, the relationship between Spm and dehydrins has not been fully elucidated. Spm, one of the most active PAs, was suggested to acts as a signaling regulator during stress. Spm Cabozantinib regulated the generation of nitric oxide signal in Arabidopsis thaliana seedlings and also interacted with ethylene or ABA to improve drought tolerance of plants. In blueberry, changes in dehydrins expression depended on endogenous ABA levels and drought intensity. Vaseva et al. reported dehydrins along with higher genes transcript levels encoding dehydrins than the sensitive one when they were subjected to drought stress. In Dendrobium candidum, the expressions of heatstable proteins and dehydrins induced by ABA have positive effects on dehydration and freezing tolerance. The similar results were carried out in Solanum species about functions of heat-stable proteins and dehydrins. We noticed exogenous Spm induced an additional heat-stable protein band about 66 kDa after 10 d of drought stress in both cultivars. Moreover, Spm significantly enhanced the accumulation of dehydrins and the transcript level of three genes encoding dehydrins in both cultivars during drought stress. These suggest that Spm is concerned with regulation of dehydrins in white clover. In addition, the data in this study also showed that drought-susceptible white clover cv. ��Ladino�� accumulated the special dehydrin after 5 d of drought, but it wasn��t observed in drought-resistant cv. ��Haifa��. Spm-induced transcript levels of dehydrin genes increased in cv. ��Ladino�� earlier than that in cv. ��Haifa��. Thus, it could be revealed that the synthesis of dehydrins and genes expression encoding dehydrins regulated by exogenously applied Spm are correlated with white clover cultivars with different drought tolerance. The studies of Blackman et al. and Walters et al. suggested that interactions between sugars and heat-stable proteins improved the dehydration tolerance of plants. However, the correlation between carbohydrate and dehydrins related to Spm regulation can��t be fully elucidated in this study and deserves further investigation. The triglyceride lipase gene subfamily is comprised of three evolutionarily related lipases: lipoprotein lipase, hepatic lipase, and endothelial lipase, and plays a central role in plasma lipoprotein metabolism and homeostasis. These lipases are differentiated by their tissue-specific gene expression, and substrate specificity. LPL is mainly expressed in adipose and muscle tissues, while HL is specifically expressed in the liver. In contrast, EL is a newly identified lipase that is synthesized by vascular endothelial cells, thyroid epithelial cells, and hepatocytes.